India stuttered in what should have been a comfortable run chase before Harmanpreet took India home with an unbeaten 31 of 32 balls.
Sri Lanka squandered a fine start to post a below par 125 for seven.
The third and final T20 will be played here on Monday.
It was a day to remember for Mandhana who became the second fastest Indian woman to reach 2,000 runs T20 runs.
Mandhana reached the feat in her 84th innings, becoming the third Indian woman after the legendary Mithali Raj (70 innings) and current skipper Harmanpreet Kaur (88 innings) to cross the mark.
Opting to bat first and with the series on line, it was the duo of captain Chamari Athapaththu (43 off 41 balls) and Vishmi Gunaratne (45 off 50 balls) provided an ideal start for the Lankans by taking the Indian bowlers to the cleaners.
The duo also put up Sri Lanka’s best opening stand (87) in T20s to leave the Indians frantically rummaging for breakthroughs.
However, as both Athapaththu and Gunaratne departed, it was a sorry state of affairs for Sri Lanka as they lost a slew of wickets, only managing an underwhelming total.
Deepti Sharma (2/34 in 4 overs) was certainly the best bowler on view while the likes of Radha Yadav and Pooja Vastrakar also helped India claw back with the ball.
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